Gaining local accuracy while not losing generality - Extending the range of gap model applications

For the study of long-term processes in forests, gap models generally sacrifice accuracy (i.e., simulating system behavior in a quantitatively accurate manner) for generality (i.e., representing a broad range of systems' behaviors with the same model). We selected the gap model ForClim to evaluate whether the local accuracy of forest succession models can be increased based on a parsimonious modeling approach that avoids the additional complexity of a 3D crown model, thus keeping parameter requirements low. We improved the representation of tree crowns by introducing feedbacks between (i) light availability and leaf area per tree and (ii) leaf area per tree and diameter growth rate to account for the self-pruning in real stands. The local accuracy of the new model, ForClim v2.9.5, was considerably improved in simulations at three long-term forest research sites in the Swiss Alps, while its generality was maintained as shown in simulations of potential natural vegetation along a broad, environmental gradient in Central Europe. We conclude that the predictive ability of a model does not depend on its complexity, but on the reproduction of patterns. Most importantly, model complexity should be consistent with the objectives of the study and the level of system understanding.