A gated channel into the proteasome core particle

Michael Groll, Monica Bajorek, Alwin Köhler, Luis Moroder, David M. Rubin, Robert Huber, Michael H. Glickman, Daniel Finley

Research output: Contribution to journalArticlepeer-review

696 Scopus citations

Abstract

The core particle (CP) of the yeast proteasome is composed of four heptameric rings of subunits arranged in a hollow, barrel-like structure. We report that the CP is autoinhibited by the N-terminal tails of the outer (α) ring subunits. Crystallographic analysis showed that deletion of the tail of the α3-subunit opens a channel into the proteolytically active interior chamber of the CP, thus derepressing peptide hydrolysis. In the latent state of the particle, the tails prevent substrate entry by imposing topological closure on the CP. Inhibition by the α-subunit tails is relieved upon binding of the regulatory particle to the CP to form the proteasome holoenzyme.

Original languageEnglish
Pages (from-to)1062-1067
Number of pages6
JournalNature Structural Biology
Volume7
Issue number11
DOIs
StatePublished - 2000
Externally publishedYes

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